The Guardian Knights are ancient warriors trapped under the ground for 600 years. These warriors are very mysterious, and the ones we know of are Commander Honcho, Guardian Willow, Dingy (the traitor) and Guardian Crackle Knuckle. Their biggest enemy is the Shadow Vespid.
Dingy was the first knight found by Loch, claiming to be the Commander of the Knights. Eventually he is controlled by Ka, and he betrayes his fellow Guardian Knights. Willow stays above ground and trains Cadets, while Crackle Knuckle keeps his bombs ready. And Honcho is waiting, waiting, for new Guardian Knights. Will you be Ook's new Guardian Knight, or should I say, hero?